Арборея

You must choose either to place one Villager from your supply onto a Pilgrimage track or advance one Pilgrimage track one space; you may pay Spirit to do one of these options a second time on your turn (p6).

When a track moves, players may take any number of their Villagers off that track and place them at the start of any trail the track has passed or is beside, but Villagers placed this way do nothing until you activate them during Step 2 of your turn (pp6–7).

Water is a wild Biome but you may only use it as another Biome type if the Biome you are replacing is at 0 on the Biome Reserve (p7).

After your turn finishes (during the next player's turn) you add completed Ecosystem cards first, then place any Invited Creatures into your Ecosystem on empty intersections not orthogonally adjacent to other Creatures; any Invited Creatures you don't place must go into Captivity, and each Creature remaining in Captivity costs you 3 Spirit at game end (pp9,12).

You may activate up to two Villagers for free during Step 2; activating a third Villager in the same turn costs you 2 Spirit (p7).

When the Sun Tracker reaches the final space the current player finishes their turn and takes the two stacked Sun Tokens; after that player finishes two more complete turns (discarding one Sun token after the first and the second after the next), you proceed to endgame scoring (p12).